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Scrats epic pursuit of his elusive acorn catapults him outside of Earth, where he accidentally sets off a series of cosmic events that transform and threaten the planet. To save themselves from peril, Manny, Sid, Diego, and the rest of the herd leave their home and embark on a quest full of thrills and spills, highs and lows, laughter and adventure while traveling to exotic new lands and encountering a host of colorful new characters.
Scrat is once again trying to bury his acorn but accidentally activates an abandoned alien ship that takes him into deep space, where he unwittingly sends several asteroids en route to a collision with Earth. Meanwhile, Manny and Ellie are preparing upcoming marriage between Peaches and her fiance, Julian. Diego and his wife Shira want to start a family, but their fierce appearance tends to scare kids. Sid is dumped by his girlfriend, Francine, just as he is about to propose to her, and he laments his solitude. During Manny and Ellies wedding anniversary party, some of the asteroids strike the place and The Herd barely escape. Meanwhile, at the underground lost world, Buck returns a dinosaur egg back to its rightful owner after it was stolen by a trio of flying tyrannosaurs named Gavin, Gertie, and Roger. Buck discovers an ancient stone pillar and takes it to the surface, where he meets Manny and the others..
Ray Romano
Manny
Denis Leary
Diego
John Leguizamo
Sid
Chris Wedge
Scrat
Stephanie Beatriz
Gertie
Robert Cardone
Freaky Bobby
Neil deGrasse Tyson
Neil deBuck Weasel
Adam Devine
Julian
Jesse Tyler Ferguson
Shangri Llama
Max Greenfield
Roger
Jessie J
Brooke
Queen Latifah
Ellie
Jennifer Lopez
Shira
Jorge Lucas
Grandpa Molehog
Andrew Christopher Nichols
Andrew Aardvark
Nick Offerman
Gavin
Keke Palmer
Peaches
James Palumbo
Panicked Start
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